箭竹-岷江冷杉林是川西亚高山暗针叶林的重要类型,研究其优势树种的空间分布格局和空间关联性对其恢复和保育具有参考意义。采用林木定位调查法,分别用Ripley’sL(t)和L12(t)函数分析群落中优势树种的空间分布格局和空间关联。结果表明:群落的优势树种为岷江冷杉(Abies faxoniana)、红桦(Betula albosinensis)和云杉(Picea asperata),群落层次结构分明。3个优势树种在小尺度上均为聚集分布,其分布格局随尺度增大而变化。岷江冷杉种群在0~11m的尺度上呈聚集分布,而在11~20m的尺度上呈随机分布;红桦种群在0~14m的尺度上呈聚集分布,而在14~20m的尺度上呈随机分布;云杉种群在0~20m尺度上均呈聚集分布。3个优势树种间的空间关联性在0~20m尺度上均呈显著的相互独立关系,表明它们相互之间作用微弱。
Bamboo-faxon fir forest is one of the most important dark coniferous forests in western Sichuan province,China.The spatial patterns and associations of dominant tree species in the forest were researched to offer references for restoration and conservation of the dark coniferous forests.Using Ripley’s univariate L(t)and bivariate L12(t) functions,we analyzed the spatial patterns and inter-specific associations of three dominant tree species in a 40 m×40 m primeval bamboo-faxon fir forest plot in sub-alpine area of western Sichuan,China.The results showed that stratification of the vertical structure of the plant community was obvious.There were three dominant tree species which Abies faxoniana,Betula albosinensis and Picea asperata.L(t).Function analysis showed the spatial patterns of three dominant tree species were changed with scales.A.faxoniana was significantly aggregated at 0~11 m scale,while randomly distributed at 11~20 m scale.B.albosinensiswas significantly aggregated at 0~14 m scale,while randomly distributed at 14~20 m scale.P.asperata clumped and was distributed at 0~20 m scale.From the analysis of L12(t)function,the results showed three dominant tree species were significantly independent with each other at the scales of 0~20 m,indicating that those species had little influence on each other.
[1] Raventós J,Wiegand T,Luis M D.Evidence for the spatial segregation hypothesis:A test with nine-year survivorship data in a Mediterranean shrubland[J].Ecology,2010,91(7):2110-2120.
[2] Das A,Battles J,Mantgem P J v,et al.Spatial elements of mortality risk in old-growth forests[J].Ecology,2008,89(6):1744-1756.
[3] Miriti M N.Twenty years of changes in spatial association and community structure among desert perennials[J].Ecology,2007,88(5):1177-1190.
[4] Dale M R T.Spatial pattern analysis in plant ecology[M].London:Cambridge University Press,1999.
[5] 刘世荣,史作民,马姜明,等.长江上游退化天然林恢复重建的生态对策[J].林业科学,2009,45(2):120-124.
[6] 四川植被协作组.四川植被[M].成都:四川人民出版社,1980.
[7] 四川森林编辑委员会.四川森林[M].北京:中国林业出版社,1992.
[8] Taylor A H,Jiang Shiwen,Zhao Lianjun,et al.Regeneration patterns and tree species coexistence in old-growth Abies-Picea forests in southwestern China[J].Forest Ecology and Management,2006,223:303-317.
[9] Taylor A H,Qin Zisheng.Tree replacement patterns in subalpine Abies-Betula forests,Wolong Natural Reserve,China[J].Plant Ecology,1988,78(3):141-149.
[10] Taylor A H,Qin Zisheng.Regeneration patterns in old-growth Abies-Betula forests in the Wolong Natural Reserve,Sichuan,China[J].Journal of Ecology,1988,76(4):1204-1218.
[11] Taylor A H,Qin Zisheng,Liu Jie.Structure and dynamics of subalpine forests in the Wanglang Natural Reserve,Sichuan,China[J].Plant Ecology,1996,124(1):25-38.
[12] 王微,陶建平,胡凯,等.华西箭竹对岷江冷杉林主要乔木树种幼苗结构及分布格局的影响[J].林业科学,2007,43(1):1-7.
[13] 李媛,陶建平,王永健,等.亚高山暗针叶林林缘华西箭竹对岷江冷杉幼苗更新的影响[J].植物生态学报,2007,31(2):283-290.
[14] Taylor A H,Huang Jinyan,Zhou Shiqiang.Canopy tree development and undergrowth bamboo dynamics in old-growth Abies-Betula forests in southwestern China:A 12-year study[J].Forest Ecology and Management,2004,200:347-360.
[15] Taylor A H,Qin Z.Tree regeneration after bamboo die-back in Chinese Abies-Betula forests[J].Journal of Vegetation Science,1992,3(2):253-260.
[16] 赵常明,陈庆恒,乔永康.青藏高原东缘岷江冷杉天然群落的种群结构和空间分布格局[J].植物生态学报,2004,28(3):341-350.
[17] 缪宁,刘世荣,史作民,等.川西亚高山红桦-岷江冷杉林优势种群的空间格局分析[J].应用生态学报,2009,20(6):1263-1270.
[18] Dale M R T,Dixon P,Fortin M-j,et al.Conceptual and mathematical relationships among methods for spatial analysis[J].Ecography,2002,25:558-577.
[19] Illian J,Penttinen A,Stoyan H,et al.Statistical analysis and modelling of spatial point patterns[M].Chichester:John Wiley & Sons,2008.
[20] Thioulouse J,Chessel D,Dolédec S,et al.Ade-4:A multivariate analysis and graphical display software[J].Statistics and Computing,1997,7:75-83.
[21] Stoyan D,Stoyan H.Fractals,random shapes,and point fields:Methods of Geometrical Statistics[M].New York:Wiley,1994.
[22] Takahashi K,Homma K,Vetrova V P,et al.Stand structure and regeneration in a Kamchatka mixed boreal forest[J].Journal of Vegetation Science,2001,12:627-634.
[23] Wiegand T,Gunatilleke S,Gunatilleke N,et al.Analyzing the spatial structure of a Sri Lankan tree species with multiple scales of clustering[J].Ecology,2007,88(12):3088-3102.
[24] Condit R,Ashton P S,Baker P,et al.Spatial patterns in the distribution of tropical tree species[J].Science,2000,288(5470):1414-1418.
[25] 缪宁,刘世荣,史作民,等.青藏高原东缘林线杜鹃-岷江冷杉原始林的空间格局[J].生态学报,2011,31(1):1-9.
[26] Zhang J,Song B,Li B H,et al.Spatial patterns and associations of six congeneric species in an old-growth temperate forest[J].Acta Oecologica,2010,36(1):29-38.
[27] 缪宁,史作民,冯秋红,等.川西亚高山岷江冷杉种群的空间格局分析[J].林业科学,2008,44(12):1-6.
[28] Martíneza I,Wiegand T,González-Taboadab F,et al.Spatial associations among tree species in a temperate forest community in north-western Spain[J].Forest Ecology and Management,2010,260(4):456-465.
[29] Hou J H,Mi X C,Liu C R,et al.Spatial patterns and associations in a Quercus-Betula forest in northern China[J].Journal of Vegetation Science,2004,15(3):407-414.
[30] 管中天.森林生态研究与应用[M].成都:四川科学技术出版社,2005.
