森林发展类型是以森林经营的经济需求和技术可行性相结合的森林经营模式。设计未来森林目标、树种选择和经营措施是森林经营设计中亟待解决的重要技术环节。采用多物种方差比值(VR),χ2检验,Pearson相关系数和Spearman秩相关系数以及Ochiai指数,Dice指数,Jaccard指数和AC联结系数对20个人工林优势种进行种间关联分析。结果表明:种对间的极显著正相关种对数比例较小,群落正处在演替早期阶段;马尾松和红椎、大叶栎的种对间有显著正联结,正联结的种对数较少且程度相对较弱,其余种对无联结或不显著的正、负联结居多。以种间联结为指标来量化树种搭配配置,通过树种结构调整、乡土珍贵树种选择,设计科学的混交种植经营模式,以期实现多样化、高价值和稳定的人工林格局。
The interspecific associations of tree species into forest management planning models have become increasingly important in sustainable forest management.Tree selection is one of the important factors of designing forest management plans.In order to study the interspecific associations of tree species affecting the adaptation of plant communities in the process of forest management,we survey 238 sample plots’ data on Masson pine plantation forest in Guangxi Province.A series of techniques including the VR analysis,χ2-test,Pearson correlation coefficient test,Spearman rank correlation coefficient test and Ochiai were used together to analyze the interspecific associations of dominant species.The results indicated that the dominant populations had general positive interspecific association,but the species pairs with significant positive associations were only ten percent of the total 190 species pairs,and most of the species pairs showed weak interspecific association.The interactions among the tree species were generally found to be complementary,yet sometimes contradictory.It was more practical for forest management from the view of multiple interspecific association and tree species selection,instead of a single species.Our study highlights the importance of having a good understanding of the tree selection of target stand as a basis for making decisions regarding forest management.
[1] 陆元昌.近自然森林经营的理论与实践[M].北京:科学出版社,2006:249.
[2] 陆元昌,张守攻,雷相东,等.人工林近自然化改造的理论基础和实施技术[J].世界林业研究,2009,22(1):20-27.
[3] Larsen J B,Nielsen A B.Close-to-nature forestry-participatory planning and educational outreach:using Forest Development Types (FDT)in communication and learning[J].Podravski Zbornik,2011,94:43-54.
[4] 彭少麟,周厚诚,郭少聪,等.鼎湖山地带性植被种间联结变化研究[J].植物学报,1999,41(11):1239-1244.
[5] 张峰,张金屯,韩广业.历山自然保护区猪尾沟森林群落树种种间关系及环境解释[J].植物生态学报,2002,26(S1):52-56.
[6] 姜俊,赵秀海.吉林蛟河针阔混交林群落优势种群种间联结性[J].林业科学,2012,47(12):149-153.
[7] 陆元昌,刘宪钊,包源,等.多功能目标下的近自然森林经营作业法研究[J].西南林业大学学报,2011,31(4):1-6.
[8] 刘宪钊,陆元昌,周燕华,等.加勒比松人工林近自然化改造中的发展类型设计[J].南京林业大学学报:自然科学版,2009,33(5):46.
[9] 张峰,上官铁梁.山西翅果油树群落种间关系的数量分析[J].植物生态学报,2000,24(3):351-355.
[10] 李意德,许涵,陈德祥,等.从植物种群间联结性探讨生态种组与功能群划分——以尖峰岭热带低地雨林乔木层数据为例[J].林业科学,2007,43(4):9-16.
[11] 简敏菲,刘琪璟,朱笃,等.九连山常绿阔叶林乔木优势种群的种间关联性分析[J].植物生态学报,2009,33(4):672-680.
[12] 王乃江,张文辉,陆元昌.陕西子午岭森林植物群落种间联结性[J].生态学报,2010,30(1):67-78.
[13] 李婷婷,陆元昌,张显强,等.经营的马尾松森林类型发育演替阶段量化指标研究[J].北京林业大学学报,2014,36(3):9-17.
[14] 王伯荪,彭少麟.南亚热带常绿阔叶林种间联结测定技术研究——I.种间联结测式的探讨与修订[J].植物生态学与地植物学丛刊,1985,9(4):274-285.
[15] Schluter D.A variance test for detecting species association with some example applications[J].Ecology,1984,65(3):998-1005.
[16] 陆元昌,雷相东,洪玲霞,等.近自然森林经理计划技术体系研究[J].西南林学院学报,2010,30(1):1-5.
[17] 孟京辉,陆元昌,王懿祥,等.海南白沙2类退化天然次生林的森林发展类型[J].林业科学,2011,47(8):44-47.
[18] Dice L R.Measure of the amount of ecological association between species[J].Ecology,1945,26(3):297-302.
[19] O’Connor I,Aarssen L W.Species association patterns in abandoned sand quarries[J].Vegetatio,1987,73 (2):101-109.
[20] 康冰,刘世荣,蔡道雄.南亚热带人工杉木林灌木层物种组成及主要木本种间联结性[J].生态学报,2005,25(9):2173-2179.
